beautiful Fossil Tooth of a Woolly Mammoth! Mammoth species can be identified from the number of enamel ridges (or lamellar plates) on their molars; primitive species had few ridges, and the number increased gradually as new species evolved to feed on more abrasive food items. [11] American president Thomas Jefferson, who had a keen interest in palaeontology, was partially responsible for transforming the word "mammoth" from a noun describing the prehistoric elephant to an adjective describing anything of surprisingly large size. [56], The woolly mammoth was probably the most specialised member of the family Elephantidae. It was used for manipulating objects, and in social interactions. It is in these circumstances that a battle of ownership occurs.. [122] It has been proposed that these changes are consistent with the concept of genomic meltdown;[121] however, the sudden disappearance of an apparently stable population may be more consistent with a catastrophic event, possibly related to climate (such as icing of the snowpack) or a human hunting expedition. [71], The best-preserved head of a frozen adult specimen, that of a male nicknamed the "Yukagir mammoth", shows that woolly mammoths had temporal glands between the ear and the eye. [119] The population seems to have subsequently been stable, without suffering further significant loss of genetic diversity. Please refer to the appropriate style manual or other sources if you have any questions. [77], The habitat of the woolly mammoth is known as "mammoth steppe" or "tundra steppe". [46] A 2011 study showed that light individuals would have been rare. This feature may have helped the mammoths to live at high latitudes. [78] The Altai-Sayan assemblages are the modern biomes most similar to the "mammoth steppe". A January Fossil of the Month. Teeth range in size from about an inch at birth to 9-12 inches in the sixth and final set. [90], Woolly mammoth bones were used as construction material for dwellings by both Neanderthals and modern humans during the ice age. Encyclopaedia Britannica's editors oversee subject areas in which they have extensive knowledge, whether from years of experience gained by working on that content or via study for an advanced degree. [45], Preserved woolly mammoth fur is orange-brown, but this is believed to be an artefact from the bleaching of pigment during burial. The appearance and behaviour of this species are among the best studied of any prehistoric animal because of the discovery of frozen carcasses in Siberia and North America, as well as skeletons, teeth, stomach contents, dung, and depiction from life in prehistoric cave paintings. [26], Since many remains of each species of mammoth are known from several localities, reconstructing the evolutionary history of the genus through morphological studies is possible. [10] It may be a version of mehemot, the Arabic version of the biblical word "behemoth". [133], In 1977, the well-preserved carcass of a seven- to eight-month-old woolly mammoth calf named "Dima" was discovered. The "Berezovka mammoth" during excavation in 1901 (left), and a model partially covered by its skin, "Dima", a frozen calf, during excavation (left), and as exhibited in the Museum of Zoology; note fur on the legs, The frozen calf "Yuka" (left), and its skull and jaw which may have been extracted from the carcass by prehistoric humans, Models of an adult and the calf "Dima" in, Mol, D. et al. Males reached shoulder heights between 2.7 and 3.4m (8.9 and 11.2ft) and weighed up to 6 metric tons (6.6 short tons). Corrections? In 1864, douard Lartet found an engraving of a woolly mammoth on a piece of mammoth ivory in the Abri de la Madeleine cave in Dordogne, France. The maturity of this ingested vegetation places the time of death in autumn rather than in spring, when flowers would be expected. By about 100,000 to 200,000 years ago, North America was home to at least two main types of mammoths: woolly mammoths in the north, and Columbian mammoths as far south as Mexico. [183] In 1899, Henry Tukeman detailed his killing of a mammoth in Alaska and his subsequent donation of the specimen to the Smithsonian Institution in Washington, DC. Two alleles were found: a dominant (fully active) and a recessive (partially active) one. They grew between eight and 11 feet tall and could weigh approximately 13,000. [28], Individuals and populations showing transitional morphologies between each of the mammoth species are known, and primitive and derived species coexisted until the former disappeared. Females averaged 2.6-2.9 m (8.5-9.5 ft) in height and weighed up to 4 tons (4.4 short tons). The leg bone once belonged to a Columbian mammoth, a short-haired elephant-like creature that wandered Florida during the Pleistocene era between 2.6 million and 10,000 years ago. This adult male specimen was called the "Yukagir mammoth", and is estimated to have lived around 18,560 years ago, and to have been 282.9cm (9.2ft) tall at the shoulder, and weighed between 4 and 5 tonnes. "This DNA is incredibly old. When it comes to a woolly mammoth vs mastodon, woolly mammoths were taller and heavier. [84] Recent stable isotope studies of Siberian and New World mammoths have shown there were differences in climatic conditions on either side of the Bering land bridge (Beringia), with Siberia being more uniformly cold and dry throughout the Late Pleistocene. Some accumulations are thought to be the remains of herds that died together at the same time, perhaps due to flooding. A 2019 study found that woolly mammoth ivory was the most suitable bony material for the production of big game projectile points during the Late Plesistocene. University of Michigan Professor Dan Fisher has been leading the dig to remove the mammoth's remains from Bristle's property this week. Several alterations in circadian clock genes were found, perhaps needed to cope with the extreme polar variation in length of daylight. The teeth had up to 26 separated ridges of enamel, which were themselves covered in "prisms" that were directed towards the chewing surface. The reason for the smaller size is unknown. where was glenn b anderson born; where did the raiders name come from; how to wire 3 phase. About 23cm (9.1in) of the crown was within the jaw, and 2.5cm (1in) was above. R538 Size: Hair Sample in a 3" x 4" zip lock bag 314). [80], The southernmost woolly mammoth specimen known is from the Shandong province of China, and is 33,000 years old. The analysis showed that the woolly mammoth and the African elephant are 98.55% to 99.40% identical. Woolly mammoths may have used their tusks as shovels to clear snow from the ground and reach the vegetation buried below, and to break ice to drink. [82][83] DNA studies have helped determine the phylogeography of the woolly mammoth. [138] While in Yakutsk in 1806, Michael Friedrich Adams heard about the frozen mammoth. [177], Local dealers estimate that 10 million mammoths are still frozen in Siberia, and conservationists have suggested that this could help save the living species of elephants from extinction. [86], A 2008 genetic study showed that some of the woolly mammoths that entered North America through the Bering land bridge from Asia migrated back about 300,000 years ago and had replaced the previous Asian population by about 40,000 years ago, not long before the entire species became extinct. [149] "Lyuba" is believed to have been suffocated by mud in a river that its herd was crossing. Several Venus figurines, including the Venus of Brassempouy and the Venus of Lespugue, were made from this material. Like their thick coat of fur, their shortened . The 10-inch-long brown, black and beige chomper, broken in two and missing a chunk, once belonged to a woolly mammoth, an elephantine creature that roamed the grassy valley that's now San. The woolly mammoth began to diverge from the steppe mammoth about 800,000 years ago in East Asia. It had long, curved tusks and four molars, which were replaced six times during the lifetime of an individual. Regional and intermediate species and subspecies such as M. intermedius, M. chosaricus, M. p. primigenius, M. p. jatzkovi, M. p. sibiricus, M. p. fraasi, M. p. leith-adamsi, M. p. hydruntinus, M. p. astensis, M. p. americanus, M. p. compressus and M. p. alaskensis have been proposed. [6], In 1796, French biologist Georges Cuvier was the first to identify the woolly mammoth remains not as modern elephants transported to the Arctic, but as an entirely new species. Woolly mammoths had broad flaps of skin under their tails which covered the anus; this is also seen in modern elephants. [137] In more recent years, scientific expeditions have been devoted to finding carcasses instead of relying solely on chance encounters. The expansion identified on the trunk of "Yuka" and other specimens was suggested to function as a "fur mitten"; the trunk tip was not covered in fur, but was used for foraging during winter, and could have been heated by curling it into the expansion. For a tooth of that quality, about $10 a lb. [115], The decline of the woolly mammoth could have increased temperatures by up to 0.2C (0.36F) at high latitudes in the Northern Hemisphere. However, at the end of the late Pleistocene about 12,000 years ago, these "megafauna" went extinct, a die-off called the Quaternary extinction. Woolly mammoths stood about 3 to 3.7 metres (about 10 to 12 feet) tall and weighed between 5,500 and 7,300 kg (between about 6 and 8 tons). Because the species was social and gregarious, creating a few specimens would not be ideal. [134][135], By 1929, the remains of 34 mammoths with frozen soft tissues (skin, flesh, or organs) had been documented. Picture 1 of 8. The coloration is a result of vivianite growing on the tusk, which. [13][29][30], A 2011 genetic study showed that two examined specimens of the Columbian mammoth were grouped within a subclade of woolly mammoths. [169][170] Woolly mammoth tusks had been articles of trade in Asia long before Europeans became acquainted with them. In addition to the technical problems, not much habitat is left that would be suitable for elephant-mammoth hybrids. [39] The well-preserved trunk of a juvenile specimen nicknamed "Yuka" was described in 2015, and it was shown to possess a fleshy expansion a third above the tip. At the time of writing, the highest bid was $7,300 (more than 5.5 lakh). Some of the bones used for materials may have come from mammoths killed by humans, but the state of the bones, and the fact that bones used to build a single dwelling varied by several thousands of years in age, suggests that they were collected remains of long-dead animals. Some cave paintings show woolly mammoths with small or no tusks, but whether this reflected reality or was artistic license is unknown. Mammoths are not elephants. All. Woolly mammoths were largely extinct by about 10,000 years ago, due to the pressures of a warming climate (which reduced the habitat of these cold-adapted mammals) combined with hunting by humans. [55] Trackways made by a woolly mammoth herd 11,30011,000 years ago have been found in the St. Mary Reservoir in Canada, showing that in this case almost equal numbers of adults, subadults, and juveniles were found. [52][50], Woolly mammoths had four functional molar teeth at a timetwo in the upper jaw and two in the lower. 8. Cave paintings of woolly mammoths exist in several styles and sizes. The diet of the woolly mammoth was mainly grasses and sedges. It features a faint reddish-brown body with dark-colored fur covering it. Rather than oval as the rest of the trunk, this part was ellipsoidal in cross section, and double the size in diameter. The glands are used especially by males to produce an oily substance with a strong smell called temporin. This is indicated on many preserved tusks by flat, polished sections up to 30 centimetres (12in) long, as well as scratches, on the part of the surface that would have reached the ground (especially at their outer curvature). In 1999, this 20,380-year-old carcass and 25 tons of surrounding sediment were transported by an Mi-26 heavy lift helicopter to an ice cave in Khatanga. [183] Due to the large area of Siberia, the possibility that woolly mammoths survived into more recent times cannot be completely ruled out, but evidence indicates that they became extinct thousands of years ago. Wooly Mammoth Tooth $375.00. Some of the hairs on . "Complete Columbian mammoth mitogenome suggests interbreeding with woolly mammoths", "Million-year-old DNA sheds light on the genomic history of mammoths", "Million-year-old mammoth genomes shatter record for oldest ancient DNA", "Collection of radiocarbon dates on the mammoths (, "Nuclear Gene Indicates Coat-Color Polymorphism in Mammoths", "Megafaunal split ends: microscopical characterisation of hair structure and function in extinct woolly mammoth and woolly rhino", "Elephantid genomes reveal the molecular bases of Woolly Mammoth adaptations to the arctic", "Mammoth Genomes Provide Recipe for Creating Arctic Elephants", "Signals of positive selection in mitochondrial proteincoding genes of woolly mammoth: Adaptation to extreme environments? He discussed the question of whether or not the remains were from elephants, but drew no conclusions. How much is a woolly mammoth tooth worth? [87] Fossils of woolly mammoths and Columbian mammoths have been found together in a few localities of North America, including the Hot Springs sinkhole of South Dakota where their regions overlapped. The owner of the real estate can argue that she is in constructive possession of the treasure, as it was located on her land. [56] A 2021 study indicates, however, that although humans likely exerted a significant selective pressure on mammoths that led to them going extinct earlier than they otherwise would have,[131] the final impetus for mammoth extinction was likely vegetation changes caused by a changed precipitation regime at the end of the Ice Age. In 2016, a group of researchers genetically examined a sample of the meal, and found it to belong to a green sea turtle (it had also been claimed to belong to Megatherium). The expansion could be used to melt snow if a shortage of water to drink existed, as melting it directly inside the mouth could disturb the thermal balance of the animal. Many are certainly known to have been killed in rivers, perhaps through being swept away by floods. A fisherman who reeled in a woolly mammoth tooth sold it at auction for more . The small ears reduced heat loss and frostbite, and the tail was short for the same reason, only 36cm (14in) long in the "Berezovka mammoth". Mastodons usually didn't grow to be over 10 ft tall, and they weighed between 4 to 6 tons. The woolly mammoth likely moulted seasonally, and the heaviest fur was shed during spring. Courtesy The Inn at Honey Run. Mammoth & Mastodon Shark Teeth By Species. [64], In 2012, a juvenile was found in Siberia, which had man-made cut marks. SHELDON, Iowa (KCAU) A woolly mammoth tooth was found in early March on the property owned by Northwest Iowa Community College (NCC) in Sheldon. It is one of the best-preserved mammoths ever found due to the almost complete head, covered in skin, but without the trunk. The closest known relatives of the Proboscidea are the sirenians (dugongs and manatees) and the hyraxes (an order of small, herbivorous mammals). [2] The first woolly mammoth remains studied by European scientists were examined by Hans Sloane in 1728 and consisted of fossilised teeth and tusks from Siberia. The origin of these remains was long a matter of debate, and often explained as being remains of legendary creatures. [163], Some researchers question the ethics of such recreation attempts. A less complete juvenile, nicknamed "Mascha", was found on the Yamal Peninsula in 1988. The chewing surface and roots are nicely preserved. [28], The first known members of the genus Mammuthus are the African species Mammuthus subplanifrons from the Pliocene, and M. africanavus from the Pleistocene. Before this, Neanderthals had co-existed with mammoths during the Middle Palaeolithic and already used mammoth bones for tool-making and building materials. The group that became extinct earlier stayed in the middle of the high Arctic, while the group with the later extinction had a much wider range. (2001). [179], Stories abound about frozen woolly mammoth meat that was consumed once defrosted, especially that of the "Berezovka mammoth", but most of these are considered dubious. $0.01 + $55.00 shipping. Woolly mammoths were around 13 feet (4 meters) tall and weighed around 6 tons (5.44 metric tons), according to the International Union for Conservation of Nature (IUCN). [140][141], The 1901 excavation of the "Berezovka mammoth" is the best documented of the early finds. 3. I know that it is pretty much universally hated by the fandom, but the designs from the 2013 walking with dinosaurs movie were very accurate for the time. Mammoth. This tooth is a manageable size for most collectors at 5-1/4" x 4-1/2 straight line measurement. [92], Woolly mammoth ivory was used to create art objects. The species is named for the appearance of its long thick coat of fur. They had a yellowish brown undercoat about 2.5 cm (about 1 inch) thick beneath a coarser outer covering of dark brown hair that grew more than 70 cm (27.5 inches) long in some individuals. These were quite wear-resistant and kept together by cementum and dentine. The cell would then be stimulated into dividing and inserted back into a female elephant. A newborn calf weighed about 90 kilograms (200 lb). Resolutions to historical issues about the validity of the genus name Mammuthus and the type species designation of E. primigenius were also proposed. [1] Distinguishing and determining these intermediate forms has been called one of the most long-lasting and complicated problems in Quaternary palaeontology. Several carcasses have been lost because they were not reported, and one was fed to dogs. It weighs a whopping 11.2 pounds and is nearly a foot long. Get a Britannica Premium subscription and gain access to exclusive content. Root is fully intact - very rare. Its behaviour was similar to that of modern elephants, and it used its tusks and trunk for manipulating objects, fighting, and foraging. James St. John / Flickr / CC BY 2.0. [32], In 2021, DNA older than a million years was sequenced for the first time, from two mammoth teeth of Early Pleistocene age found in eastern Siberia. [119], Before their extinction, the Wrangel Island mammoths had accumulated numerous genetic defects due to their small population; in particular, a number of genes for olfactory receptors and urinary proteins became nonfunctional, possibly because they had lost their selective value on the island environment. [132], Woolly mammoth fossils have been found in many different types of deposits, including former rivers and lakes, and in "Doggerland" in the North Sea, which was dry at times during the ice age. Woolly mammoths sustained themselves on plant food, mainly grasses and sedges, which were supplemented with herbaceous plants, flowering plants, shrubs, mosses, and tree matter. The population of woolly mammoths declined at the end of the Pleistocene, disappearing throughout most of its mainland range, although isolated populations survived on St. Paul Island until 5,600 years ago, on Wrangel Island until 4,000 years ago, and possibly (based on ancient eDNA) in the Yukon up to 5,700 years ago and on the Taymyr Peninsula up to 3,900 years ago. The woolly mammoth tusk was discovered in 2017 and although valuable, the rare blue coloring makes it an exquisite piece. Today, more than 500 depictions of woolly mammoths are known, in media ranging from cave paintings and engravings on the walls of 46 caves in Russia, France, and Spain to engravings and sculptures (termed "portable art") made from ivory, antler, stone and bone. Many mammoth carcasses may have been scavenged by humans rather than hunted. World's oldest DNA discovered in 1.2-million-year-old mammoth teeth. Woolly Mammoth Fossil tooth with roots. $1,495.00. This article was most recently revised and updated by, https://www.britannica.com/animal/woolly-mammoth. Cloning would involve removal of the DNA-containing nucleus of the egg cell of a female elephant and replacement with a nucleus from woolly mammoth tissue. Picture Information. Impressive 10 Pound (4.7 KG) Woolly Mammoth Fossil Tooth Found In Siberia $1,400.00 Free shipping or Best Offer 2 Big Woolly Rhinoceros Fossil Tooth + Roots Omsk Siberia Pleistocene Ice Age Kk $119.00 $14.95 shipping or Best Offer 22" Fossil Woolly Mammoth Tibia Bone 13lb Authentic Ancient Pre-historic OLD $609.99 or Best Offer 20 watching An EXTRA LARGE, incredibly preserved Woolly Mammoth (Mammuthus primigenius), an early elephant, molar found in the Dogger Bank, North Sea. The Woolly Mammoth can beg as a pre-teen and jump as a teen. We offer genuine mammoth tusks, chunks and pieces of the prehistoric ivory and bone from Alaska, the Yukon and Siberia. The relative abundance and, at times, excellent preservation of carcasses of thisspeciesfound in thepermafrost (permanently frozen ground)of Siberia have provided much information about mammoths structure and habits. The two-fingered tip of the trunk was probably adapted for picking up the short grasses of the last ice age (Quaternary glaciation, 2.58 million years ago to present) by wrapping around them, whereas modern elephants curl their trunks around the longer grass of their tropical environments. Thriving during the Pleistocene ice ages, woolly mammoths died out after much of their habitat was lost as Earths climate warmed in the aftermath of the last ice age. The name mastodon literally means "breast tooth," referring to the the "nipple"-shaped bumps along the top edges of these animals' teeth. [8] In 1828, the British naturalist Joshua Brookes used the name Mammuthus borealis for woolly mammoth fossils in his collection that he put up for sale, thereby coining a new genus name. This habitat was not dominated by ice and snow, as is popularly believed, since these regions are thought to have been high-pressure areas at the time. Later woolly and Columbian mammoths also interbred occasionally, and mammoth species may have hybridised routinely when brought together by glacial expansion. Its cousin the Steppe mammoth ( M. trogontherii) was perhaps the largest one in the family growing up to 13 to 15 feet tall. How much is a mammoth tusk worth? The woolly mammoth has been mostly extinct for 10,000 years, with the final vestigial populations surviving until about 4,000 years ago. Both molars were thought lost by the 1980s, and the more complete "Taimyr mammoth" found in Siberia in 1948 was therefore proposed as the neotype specimen in 1990. The error was not corrected until 1899, and the correct placement of mammoth tusks was still a matter of debate into the 20th century. [17] The following cladogram shows the placement of the genus Mammuthus among other proboscideans, based on characteristics of the hyoid bone in the neck:[18] According to the Jacksonville Zoo, the woolly mammoth lived in North America and Asia until about 4,000 years ago. [181] In 2011, the Chinese palaeontologist Lida Xing livestreamed while eating meat from a Siberian mammoth leg (thoroughly cooked and flavoured with salt) and told his audience it tasted bad and like soil. Only its molars are known, which show that it had 810 enamel ridges. The fact that sperm cells of modern mammals are viable for 15 years at most after deep-freezing makes this method unfeasible. The Columbian mammoth (Mammuthus columbi) lived alongside the woolly mammoth in North America, and DNA studies show that the two hybridised with each other. When inserted into human cells, the mammoth's version of the protein was found to be less sensitive to heat than the elephant's. The woolly mammoth, scientific name Mammuthus primigenius, is related to the modern African and Asian elephants. The woolly mammoth (Mammuthis primigenius) evolved later, as the climate cooled, and was a grazer. [39], Other characteristic features depicted in cave paintings include a large, high, single-domed head and a sloping back with a high shoulder hump; this shape resulted from the spinous processes of the back vertebrae decreasing in length from front to rear. The bases of the huts were circular, and ranged from 8 to 24 square metres (86 to 258sqft). Males stood between nine and 11 feet high at the shoulder and females were slightly smaller8.5-9.5 feet tall at the shoulder. This environment stretched across northern Asia, many parts of Europe, and the northern part of North America during the last ice age. It' DNA has been successfully sequenced so an ancient woolly rhino could be created in a similar way to a mammoth. A woolly mammoth tooth found off the coast of Newburyport, Mass., sold at auction for more than $10,000. It was similar to the grassy steppes of modern Russia, but the flora was more diverse, abundant, and grew faster. The habitat of the woolly mammoth supported other grazing herbivores such as the woolly rhinoceros, wild horses, and bison. In the 19th century, several reports of "large shaggy beasts" were passed on to the Russian authorities by Siberian tribesmen, but no scientific proof ever surfaced. It was 34 months old, and a laceration on its right foot may have been the cause of death. The first molars were about the size of those of a human, 1.3cm (0.51in), the third were 15cm (6in) 15cm (5.9in) long, and the sixth were about 30cm (1ft) long and weighed 1.8kg (4lb). [79] A 2014 study concluded that forbs (a group of herbaceous plants) were more important in the steppe-tundra than previously acknowledged, and that it was a primary food source for the ice-age megafauna. [78], Modern humans co-existed with woolly mammoths during the Upper Palaeolithic period when the humans entered Europe from Africa between 30,000 and 40,000 years ago. [137] While frozen woolly mammoth carcasses had been excavated by Europeans as early as 1728, the first fully documented specimen was discovered near the delta of the Lena River in 1799 by Ossip Schumachov, a Siberian hunter. [44] Woolly mammoths had numerous sebaceous glands in their skin, which secreted oils into their hair; this would have improved the wool's insulation, repelled water, and given the fur a glossy sheen. Remains of various extinct elephants were known by Europeans for centuries, but were generally interpreted, based on biblical accounts, as the remains of legendary creatures such as behemoths or giants. [73], Evidence of several different bone diseases has been found in woolly mammoths. This tooth is suspected to be over 20,000 years old. Can scientists bring mammoths back to life by cloning? As massive as they were13 feet long and five to seven tonswoolly mammoths figured on the lunch menu of early Homo sapiens, who coveted them for their warm pelts (one of which could have kept an entire family comfy on bitterly cold nights) as well as their tasty, fatty meat. [15] The paralectotype molar (specimen GZG.V.010.018) has since been located in the Gttingen University collection, identified by comparing it with Osborn's illustration of a cast. After several generations of cross-breeding these hybrids, an almost pure woolly mammoth would be produced. Mammoths born with at least one copy of the dominant allele would have had dark coats, while those with two copies of the recessive allele would have had light coats. The woolly mammoth (Mammuthus primigenius) is an extinct species of mammoth that lived during the Pleistocene until its extinction in the Holocene epoch. The trunk could be used for pulling off large grass tufts, delicately picking buds and flowers, and tearing off leaves and branches where trees and shrubs were present. The first Siberian ivory to reach western Europe was brought to London in 1611. Large male Such remains are mostly found above the Arctic Circle, in permafrost. [72] This feature indicates that, like bull elephants, male woolly mammoths entered "musth", a period of heightened aggressiveness. A male woolly mammoth's shoulder height was 9 to 11 feet tall and weighed around 6 tons. Trade in fossil ivory is legal (and. "The Jarkov Mammoth: 20,000-Year-Old carcass of a Siberian woolly mammoth, Staatliches Museum fr Naturkunde Stuttgart, Musum national d'Histoire naturelle, Paris, Zoological Museum of the Zoological Institute of the Russian Academy of Sciences, State Museum of Natural History Stuttgart, "An Account of Elephants Teeth and Bones Found under Ground", "Of Fossile Teeth and Bones of Elephants. [182], There have been occasional claims that the woolly mammoth is not extinct and that small, isolated herds might survive in the vast and sparsely inhabited tundra of the Northern Hemisphere. $75.00 + $12.45 shipping. Different woolly mammoth populations did not die out simultaneously across their range, but gradually became extinct over time.
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